Nuclear Black Hole Formation in Clumpy Galaxies at High Redshift

Massive stellar clumps in high redshift galaxies interact and migrate to the center to form a bulge and exponential disk in <1 Gyr. Here we consider the fate of intermediate mass black holes (BHs) that might form by massive-star coalescence in the dense young clusters of these disk clumps. We find that the BHs move inward with the clumps and reach the inner few hundred parsecs in only a few orbit times. There they could merge into a supermassive BH by dynamical friction. The ratio of BH mass to stellar mass in the disk clumps is approximately preserved in the final ratio of BH to bulge mass. Because this ratio for individual clusters has been estimated to be ~10^{-3}, the observed BH-to-bulge mass ratio results. We also obtain a relation between BH mass and bulge velocity dispersion that is compatible with observations of present-day galaxies.Comment: 10 pages, 3 figures, accepted by Ap

Resolving Gas Dynamics in the Circumnuclear Region of a Disk Galaxy in a Cosmological Simulation

Using a hydrodynamic adaptive mesh refinement code, we simulate the growth and evolution of a galaxy, which could potentially host a supermassive black hole, within a cosmological volume. Reaching a dynamical range in excess of 10 million, the simulation follows the evolution of the gas structure from super-galactic scales all the way down to the outer edge of the accretion disk. Here, we focus on global instabilities in the self-gravitating, cold, turbulence-supported, molecular gas disk at the center of the model galaxy, which provide a natural mechanism for angular momentum transport down to sub-pc scales. The gas density profile follows a power-law scaling as r^-8/3, consistent with an analytic description of turbulence in a quasi-stationary circumnuclear disk. We analyze the properties of the disk which contribute to the instabilities, and investigate the significance of instability for the galaxy's evolution and the growth of a supermassive black hole at the center.Comment: 16 pages (includes appendix), submitted to ApJ. Figures here are at low resolution; for higher resolution version, download http://casa.colorado.edu/~levinerd/ms.pd

Reconstructing the massive black hole cosmic history through gravitational waves

The massive black holes we observe in galaxies today are the natural end-product of a complex evolutionary path, in which black holes seeded in proto-galaxies at high redshift grow through cosmic history via a sequence of mergers and accretion episodes. Electromagnetic observations probe a small subset of the population of massive black holes (namely, those that are active or those that are very close to us), but planned space-based gravitational-wave observatories such as the Laser Interferometer Space Antenna (LISA) can measure the parameters of ``electromagnetically invisible'' massive black holes out to high redshift. In this paper we introduce a Bayesian framework to analyze the information that can be gathered from a set of such measurements. Our goal is to connect a set of massive black hole binary merger observations to the underlying model of massive black hole formation. In other words, given a set of observed massive black hole coalescences, we assess what information can be extracted about the underlying massive black hole population model. For concreteness we consider ten specific models of massive black hole formation, chosen to probe four important (and largely unconstrained) aspects of the input physics used in structure formation simulations: seed formation, metallicity ``feedback'', accretion efficiency and accretion geometry. For the first time we allow for the possibility of ``model mixing'', by drawing the observed population from some combination of the ``pure'' models that have been simulated. A Bayesian analysis allows us to recover a posterior probability distribution for the ``mixing parameters'' that characterize the fractions of each model represented in the observed distribution. Our work shows that LISA has enormous potential to probe the underlying physics of structure formation.Comment: 24 pages, 16 figures, submitted to Phys. Rev.

Direct cosmological simulations of the growth of black holes and galaxies

We investigate the coupled formation and evolution of galaxies and their embedded supermassive black holes using state-of-the-art hydrodynamic simulations of cosmological structure formation. For the first time, we self-consistently follow the dark matter dynamics, radiative gas cooling, star formation, as well as black hole growth and associated feedback processes, starting directly from initial conditions appropriate for the LambdaCDM cosmology. Our modeling of the black hole physics is based on an approach we have developed in simulations of isolated galaxy mergers. Here we examine: (i) the predicted global history of black hole mass assembly (ii) the evolution of the local black hole-host mass correlations and (iii) the conditions that allow rapid growth of the first quasars, and the properties of their hosts and descendants today. We find a total black hole mass density in good agreement with observational estimates. The black hole accretion rate density peaks at lower redshift and evolves more strongly at high redshift than the star formation rate density, but the ratio of black hole to stellar mass densities shows only a moderate evolution at low redshifts. We find strong correlations between black hole masses and properties of the stellar systems, agreeing well with the measured local M_BH-sigma and M_BH -M_* relationships, but also suggesting (dependent on the mass range) a weak evolution with redshift in the normalization and the slope. Our simulations also produce massive black holes at high redshift, due to extended periods of exponential growth in regions that collapse early and exhibit strong gas inflows. These first supermassive BH systems however are not necessarily the most massive ones today, since they are often overtaken in growth by quasars that form later. (abridged)Comment: 22 pages, 17 figures, submitted to Ap

Toward precise constraints on growth of massive black holes

Growth of massive black holes (MBHs) in galactic centers comes mainly from gas accretion during their QSO/AGN phases. In this paper we apply an extended Soltan argument, connecting the local MBH mass function with the time-integral of the QSO luminosity function, to the demography of MBHs and QSOs from recent optical and X-ray surveys, and obtain robust constraints on the luminosity evolution (or mass growth history) of individual QSOs (or MBHs). We find that the luminosity evolution probably involves two phases: an initial exponentially increasing phase set by the Eddington limit and a following phase in which the luminosity declines with time as a power law (with a slope of -1.2--1.3) set by a self-similar long-term evolution of disk accretion. Neither an evolution involving only the increasing phase with a single Eddington ratio nor an exponentially declining pattern in the second phase is likely. The period of a QSO radiating at a luminosity higher than 10% of its peak value is about (2-3)x10^8 yr, during which the MBH obtains ~80% of its mass. The mass-to-energy conversion efficiency is $0.16\pm0.04 ^{+0.05}_{-0}$, with the latter error accounting for the maximum uncertainty due to Compton-thick AGNs. The expected Eddington ratios in QSOs from the constrained luminosity evolution cluster around a single value close to 0.5-1 for high-luminosity QSOs and extend to a wide range of lower values for low-luminosity ones. The Eddington ratios for high luminosity QSOs appear to conflict with those estimated from observations (~0.25) by using some virial mass estimators for MBHs in QSOs unless the estimators systematically over-estimate MBH masses by a factor of 2-4. We also infer the fraction of optically obscured QSOs ~60-80%. Further applications of the luminosity evolution of individual QSOs are also discussed.Comment: 25 pages, 13 figures, ApJ in pres

Reciprocal regulation of PKA and rac signaling

Activated G protein-coupled receptors (GPCRs) and receptor tyrosine kinases relay extracellular signals through spatial and temporal controlled kinase and GTPase entities. These enzymes are coordinated by multifunctional scaffolding proteins for precise intracellular signal processing. The cAMP-dependent protein kinase A (PKA) is the prime example for compartmentalized signal transmission downstream of distinct GPCRs. A-kinase anchoring proteins tether PKA to specific intracellular sites to ensure precision and directionality of PKA phosphorylation events. Here, we show that the Rho-GTPase Rac contains A-kinase anchoring protein properties and forms a dynamic cellular protein complex with PKA. The formation of this transient core complex depends on binary interactions with PKA subunits, cAMP levels and cellular GTP-loading accounting for bidirectional consequences on PKA and Rac downstream signaling. We show that GTP-Rac stabilizes the inactive PKA holoenzyme. However, β-adrenergic receptor-mediated activation of GTP-Rac–bound PKA routes signals to the Raf-Mek-Erk cascade, which is critically implicated in cell proliferation. We describe a further mechanism of how cAMP enhances nuclear Erk1/2 signaling: It emanates from transphosphorylation of p21-activated kinases in their evolutionary conserved kinase-activation loop through GTP-Rac compartmentalized PKA activities. Sole transphosphorylation of p21-activated kinases is not sufficient to activate Erk1/2. It requires complex formation of both kinases with GTP-Rac1 to unleash cAMP-PKA–boosted activation of Raf-Mek-Erk. Consequently GTP-Rac functions as a dual kinase-tuning scaffold that favors the PKA holoenzyme and contributes to potentiate Erk1/2 signaling. Our findings offer additional mechanistic insights how β-adrenergic receptor-controlled PKA activities enhance GTP-Rac–mediated activation of nuclear Erk1/2 signaling

The quasar M_bh - M_host relation through Cosmic Time II - Evidence for evolution from z=3 to the present age

We study the dependence of the M_bh - M_host relation on the redshift up to z=3 for a sample of 96 quasars the host galaxy luminosities of which are known. Black hole masses were estimated assuming virial equilibrium in the broad line regions (Paper I), while the host galaxy masses were inferred from their luminosities. With this data we are able to pin down the redshift dependence of the M_bh - M_host relation along 85 per cent of the Universe age. We show that, in the sampled redshift range, the M_bh - L_host relation remains nearly unchanged. Once we take into account the aging of the stellar population, we find that the M_bh / M_host ratio (Gamma) increases by a factor ~7 from z=0 to z=3. We show that Gamma evolves with z regardless of the radio loudness and of the quasar luminosity. We propose that most massive black holes, living their quasar phase at high-redshift, become extremely rare objects in host galaxies of similar mass in the Local Universe.Comment: 10 pages, 8 figures, 2 tables. Accepted for publication in MNRA

Angular Momentum and the Formation of Stars and Black Holes

The formation of compact objects like stars and black holes is strongly constrained by the requirement that nearly all of the initial angular momentum of the diffuse material from which they form must be removed or redistributed during the formation process. The mechanisms that may be involved and their implications are discussed for (1) low-mass stars, most of which probably form in binary or multiple systems; (2) massive stars, which typically form in clusters; and (3) supermassive black holes that form in galactic nuclei. It is suggested that in all cases, gravitational interactions with other stars or mass concentrations in a forming system play an important role in redistributing angular momentum and thereby enabling the formation of a compact object. If this is true, the formation of stars and black holes must be a more complex, dynamic, and chaotic process than in standard models. The gravitational interactions that redistribute angular momentum tend to couple the mass of a forming object to the mass of the system, and this may have important implications for mass ratios in binaries, the upper stellar IMF in clusters, and the masses of supermassive black holes in galaxies.Comment: Accepted by Reports on Progress in Physic

On the cosmic evolution of the scaling relations between black holes and their host galaxies: Broad Line AGN in the zCOSMOS survey

(Abriged) We report on the measurement of the rest frame K-band luminosity and total stellar mass of the hosts of 89 broad line Active Galactic Nuclei detected in the zCOSMOS survey in the redshift range 1<z<2.2. The unprecedented multiwavelength coverage of the survey field allows us to disentangle the emission of the host galaxy from that of the nuclear black hole in their Spectral Energy Distributions. We derive an estimate of black hole masses through the analysis of the broad Mg II emission lines observed in the medium-resolution spectra taken with VIMOS/VLT as part of the zCOSMOS project. We found that, as compared to the local value, the average black hole to host galaxy mass ratio appears to evolve positively with redshift, with a best fit evolution of the form (1+z)^{0.68 \pm0.12 +0.6 -0.3}, where the large asymmetric systematic errors stem from the uncertainties in the choice of IMF, in the calibration of the virial relation used to estimate BH masses and in the mean QSO SED adopted. A thorough analysis of observational biases induced by intrinsic scatter in the scaling relations reinforces the conclusion that an evolution of the MBH-M* relation must ensue for actively growing black holes at early times: either its overall normalization, or its intrinsic scatter (or both) appear to increase with redshift. This can be interpreted as signature of either a more rapid growth of supermassive black holes at high redshift, a change of structural properties of AGN hosts at earlier times, or a significant mismatch between the typical growth times of nuclear black holes and host galaxies.Comment: 47 pages, 8 figures. Accepted for publication in Ap

Modeling the cosmological co-evolution of supermassive black holes and galaxies: I. BH scaling relations and the AGN luminosity function

We model the cosmological co-evolution of galaxies and their central supermassive black holes (BHs) within a semi-analytical framework developed on the outputs of the Millennium Simulation. This model, described in detail in Croton et al. (2006) and De Lucia & Blaizot (2007), introduces a `radio mode' feedback from Active Galactic Nuclei (AGN) at the centre of X-ray emitting atmospheres in galaxy groups and clusters. Thanks to this mechanism, the model can simultaneously explain: (i) the low observed mass drop-out rate in cooling flows; (ii) the exponential cut-off in the bright end of the galaxy luminosity function; and (iii) the bulge-dominated morphologies and old stellar ages of the most massive galaxies in clusters. This paper is the first of a series in which we investigate how well this model can also reproduce the physical properties of BHs and AGN. Here we analyze the scaling relations, the fundamental plane and the mass function of BHs, and compare them with the most recent observational data. Moreover, we extend the semi-analytic model to follow the evolution of the BH mass accretion and its conversion into radiation, and compare the derived AGN bolometric luminosity function with the observed one. While we find for the most part a very good agreement between predicted and observed BH properties, the semi-analytic model underestimates the number density of luminous AGN at high redshifts, independently of the adopted Eddington factor and accretion efficiency. However, an agreement with the observations is possible within the framework of our model, provided it is assumed that the cold gas fraction accreted by BHs at high redshifts is larger than at low redshifts.Comment: 15 pages, 7 figures, MNRAS submitte